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Orphan Sienna, betrayed by fiancé Pierce and sister Chloe, rescues a wounded wolf—Alpha Axel of the werewolf clan. He claims her as his fated mate, bound by a three-brother pact with Lip and Zane. Amid war with snake-people and human persecution, Sienna loses Lip but finds family with Axel, Zane, and their children, building a new life in the forest.